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Does Your Web Experience Need to be Exactly the Same in Every Browser?

Nope.
But you should really try to make it consistent. Check this site out in different browsers (Firefox, Safari, etc.).
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Here's another example. Look at these images from Jeremy Cowart in Safari and then Firefox. This has to do with color profiles JPEG's are exported in.
http://blog.jeremycowart.com/2009/10/brandi-carli...
Wow Jay… that's whack…! that's not a good thing either.
I tried it in IE7 on my work computer. It won't give me the "Greetings, Modern Browser People" message. Smart website. =)
haha!